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HomePublications

In this section, you will be able to view all the completed, public SCALES deliverables as well as a listing of scientific publications resulting from the project. Project logo, posters, and leaflets can be downloaded from the Media Centre.

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  Publications - Sorted Ascending

By Keyword
  • beta diversity
  • biotic communities
  • biotic interactions
  • climate change
  • connectivity
  • dissemination of results
  • disturbance
  • downscaling
  • ecosystem services
  • ecosystems
  • effectiveness instruments
  • efficiency of instruments
  • environmental pressures
  • extinction debt
  • fragmentation
  • functional diversity
  • genes
  • governance
  • habitat loss
  • individuals
  • innovative instruments
  • isolation
  • management
  • matching of methods and instruments
  • participation
  • policy
  • populations
  • practical suitability of methods and policy instruments
  • resilience
  • scale-dependent impact
  • SCALES Tool
  • socio-economic driving forces
  • species area relationships
  • species richness
  • stability
  • upscaling

    • « 1 2 3 »
    Title ↓ ↑ Author(s) ↓ ↑ Year ↓ ↑ Journal ↓ ↑ URL/DOI ↓ ↑ Derived ↓ ↑ Size ↓ ↑

    Haldane's rule revisited: do hybrid females have a shorter lifespan? Survival of hybrids in a recent contact zone between two large gull species
    Neubauer G., Nowicki P., Zagalska-Neubauer M.
    2014
    Journal of Evolutionary Biology
    http://onlineli brary.wiley.com /doi/10.1111/je b.12404/abstrac t
    Yes

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    Butterfly dispersal in inhospitable matrix: rare, risky, but long-distance
    PDF download
    Nowicki P., Vrabec V., Binzenhöfer B., Feil J., Zakšek B., Hovestadt T., Settele J.
    2014
    Landscape Ecology
    http://link.spr inger.com/artic le/10.1007%2Fs1 0980-013-9971-0
    Yes
    546.07KB

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    Metapopulations of endangered Maculinea butterflies are resilient to large-scale fire
    Nowicki P., Kajzer-Bonk J., Marczyk J
    2014
    Ecohydrology
    http://onlineli brary.wiley.com /doi/10.1002/ec o.1484/full
    Yes

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    Bushes protected tortoises from lethal overheating in arid areas of Morocco
    Lagarde, F., Louzizi, T., Slimani, T., El Mouden, H., Ben Kaddour, K., Moulherat, S., Bonnet, X.
    2012
    Environmental Conservation
    http://journals .cambridge.org/ action/displayA bstract?fromPag e=online&aid...
    Yes

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    How far can a tortoise walk in open habitat before overheating? Implications for conservation
    Moulherat S., Delmas V., Slimani T., El Hassan El Mouden, Louzizi T., Lagardea F., Bonnet X.
    2014
    Journal for Nature Conservation
    http://www.scie ncedirect.com/s cience/article/ pii/S1617138113 00112X
    Yes

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    Emerging multilevel environmental governance – A case of public participation in Poland
    PDF download
    Cent J., Grodzińska-Jurczak M., Pietrzyk-Kaszyńska A.
    2014
    Journal for Nature Conservation
    http://www.scie ncedirect.com/s cience/article/ pii/S1617138113 000988
    Yes
    967.43KB

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    Selecting pseudo-absence for species distribution models: how, where and how many?
    Barbet-Massin M, Jiguet F, Albert C, Thuiller W.
    2012
    Methods in Ecology and Evolution
    http://onlineli brary.wiley.com /doi/10.1111/j. 2041-210X.2011. 00172.x/abst...
    Yes

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    From supply to social demand: a landscape-scale analysis of the water regulation service
    Cristina Quintas-Soriano, Antonio J. Castro, Marina García-Llorente, Javier Cabello, Hermelindo Castro
    2014
    Landscape Ecology
    http://link.spr inger.com/artic le/10.1007%2Fs1 0980-014-0032-0
    No

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    EU agricultural reform fails on biodiversity
    Pe'er, G., L. V. Dicks, P. Visconti, R. Arlettaz, A. Báldi, T. G. Benton, S. Collins et al.
    2014
    Science
    http://dx.doi.o rg/10.1126/scie nce.1253425
    Yes

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    scale effectiveness of governance in biodiversity conservation. National regulatory model of biodiversity policy in Poland
    PDF download
    Grodzińska-Jurczak M., Cent J., Pietrzyk-Kaszyńska A., Szentgyörgyi H.
    2014
    Report
    http://scales-p roject.net/img/ uplf/12964-02-P oland%20-%20Boo k.pdf
    Yes
    1.75MB

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    Scale sensitivity and scale effectiveness of governance in biodiversity conservation. National regulatory model of biodiversity policy in Finland
    PDF download
    Paloniemi R., Koivulehto M., Primmer E., Similä J.
    2014

    http://scales-p roject.net/img/ uplf/12965-04-F inland%20-%20Bo ok.pdf
    Yes
    2.42MB

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    Scale sensitivity and scale effectiveness of governance in biodiversity conservation. National regulatory model of biodiversity policy in GREECE
    PDF download
    Apostolopoulou E., Touloumis K., Pantis J. D.
    2014

    http://scales-p roject.net/img/ uplf/12966-05-G reece%20-%20Boo k.pdf
    Yes
    2.21MB

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    Genetic consequences of habitat fragmentation during a range expansion
    PDF download
    Stefano Mona, Nicolas Ray, Miguel Arenas, Laurent Excoffier
    2014
    Heredity
    http://www.natu re.com/hdy/jour nal/v112/n3/abs /hdy2013105a.ht ml
    Yes
    715.22KB

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    Predicting species distributions for conservation decisions
    PDF download
    Antoine Guisan, Reid Tingley, John B. Baumgartner, Ilona Naujokaitis-Lewis, Patricia R. Sutcliffe, Ayesha I. T. Tulloch, Tracey J. Regan, Lluis Brotons, Eve McDonald-Madden, Chrystal Mantyka-Pringle, Tara G. Martin, Jonathan R. Rhodes, Ramona Maggini, Samantha A. Setterfield, Jane Elith, Mark W. Schwartz, Brendan A. Wintle, Olivier Broennimann, Mike Austin, Simon Ferrier, Michael R. Kearney, Hugh P. Possingham, Yvonne M. Buckley
    2013
    Ecology Letters
    http://onlineli brary.wiley.com /doi/10.1111/el e.12189/abstrac t
    Yes
    1.41MB

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    Simulation of Ecosystem Responses to Multiple Hierarchical Disturbances: a Major Earthquake and Typhoons
    Chiang, L.-C., Y.-P. Lin, T. Huang, Schmeller D., Verburg P., Y.-L. Liu, and T.-S. Ding
    2013
    Landscape and Urban Planning

    Yes

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    An optimal sampling scheme with downscaling approach for sampling of bird species at various scale sample sites
    Yu-Pin Lin, Wei-Chih Lin, Tzung-Su Ding, Pei-Fen Lee, Schemeller D., Henle K., Tsai-Chu Wu
    2013


    Yes

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    STAR-BME: A GIS tool for spatiotemporal modeling under a knowledge synthesis framework
    Yu, Hwa-Lung, Shang-Chen Ku, and Kolovos A.
    2013
    Environmental Modeling & Software

    Yes

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    Land-use pattern modification for optimal bird\'s conservation
    Chun-Wei Huang, Yu-Pin Lin, Tzung-Su Ding, Yung-Chieh Wang, Wei-Te Hsiao
    2013


    Yes

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    Effectiveness of Nature Conservation – a case of Natura 2000 sites in Poland. W: Protected Area Management
    PDF download
    Grodzińska-Jurczak M., Strzelecka M., Kamal S., Gutowska J.

    InTech, Rijeka
    http://www.inte chopen.com/book s/protected-are a-management/ef fectiveness-...
    Yes
    243.98KB

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    Simple process-based simulators for generating spatial patterns of habitat loss and fragmentation: A review and introduction to the G-RaFFe model
    PDF download
    Pe\'er, G., G. A. Zurita, L. Schober, M. I. Bellocq, M. Strer, M. Müller, and S. Pütz
    2013
    PLoS ONE
    http://www.plos one.org/article /info%3Adoi%2F1 0.1371%2Fjourna l.pone.0064968
    Yes
    1.52MB

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    Minimum Area Requirements: what can we learn from the different ways to derive it, and how can we make it useful for biological conservation
    G. Pe’er, M.Α. Tsianou, K.W. Franz, Y.G. Matsinos, A.D. Mazaris, D. Storch, L. K, J. Verboom, M. Baguette, V.M. Stevens, and K. Henle

    Biological Conservation

    Yes

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    Bird-monitoring in Europe - a first overview of practices, motivations and aims
    PDF download
    Schmeller D.S. , Henle K., Loyau A., Besnard A., Henry P-Y.
    2012
    Nature Conservation
    http://www.pens oft.net/journal s/natureconserv ation/article/3 644/
    Yes
    1.64MB

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    Spatial aggregation and the species–area relationship across scales
    Grillia J., Azaeleb S., Banavarc J., Maritana A.
    2012
    Journal of Theoretical Biology
    http://www.scie ncedirect.com/s cience/article/ pii/S0022519312 003839
    Yes

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    Edge artefacts and lost performance in national versus continental conservation priority areas
    PDF download
    Moilanen, A., Anderson, B. J., Arponen, A., Pouzols, F. M., & Thomas, C. D
    2013
    Diversity and Distributions
    http://onlineli brary.wiley.com /doi/10.1111/dd i.12000/abstrac t
    Yes
    641.31KB

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    National responsibilities for conserving habitats – a freely scalable method
    PDF download
    Schmeller D.S. , Maier A., Evans D., Henle K.
    2012
    Nature Conservation
    http://www.pens oft.net/journal s/natureconserv ation/article/3 710/national...
    Yes
    4.07MB

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    Functional species pool framework to test for biotic effects on community assembly
    de Bello F., Price J.N., Münkemüller T., Liira J., Zobel,M., Thuiller W., Gerhold P., Götzenberger L., Lavergne S., Leps J., Zobel K., Pärtel M.
    2012
    Ecology
    http://www.esaj ournals.org/doi /abs/10.1890/11 -1394.1
    Yes

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    Mycorrhizas in the Central European flora: relationships with plant life history traits and ecology
    Hempel S., Götzenberger L., Kühn I., Michalski S.G., Rillig M.C., Zobel M., Moora M.

    Ecology
    http://www.esaj ournals.org/doi /abs/10.1890/12 -1700.1
    Yes

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    Evaluating the connectivity of a protected areas' network under the prism of global change: the efficiency of the European Natura 2000 network for four birds of prey
    PDF download
    Mazaris A.D., Papanikolaou A.D., Barbet-Massin M., Kallimanis A.S., Jiguet F., Schmeller D.S. , Pantis J.D.
    2013
    PLoS ONE
    http://www.plos one.org/article /info%3Adoi%2F1 0.1371%2Fjourna l.pone.0059640
    Yes
    461.84KB

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    Effectiveness of the Natura 2000 network to cover threatened species
    PDF download
    Trochet A., Schmeller D.S.
    2013
    Nature Conservation
    http://www.pens oft.net/journal s/natureconserv ation/article/3 626/abstract/
    Yes
    857.23KB

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    "Mind the gap!" - How well does Natura 2000 cover species of European interest?
    PDF download
    Gruber B., Evans D., Henle K., Bauch B., Schmeller D.S. , Dziock F., Henry P.-Y., Lengyel S., Margules C., Dormann C.F.
    2012
    Nature Conservation
    http://www.pens oft.net/journal s/natureconserv ation/article/3 732/abstract/
    Yes
    2.18MB

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    Mixed reproductive strategies of the Common Moorhen on a micro-scale as revealed by genetic data
    Loyau A., Schmeller D.S.
    2012
    Comptes Rendus Biologies
    http://www.ncbi .nlm.nih.gov/pu bmed/23199635
    Yes

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    Urbanisation effect on Orthoptera: which scale matters ? Insect Conservation and Diversity
    Penone C., Kerbiriou C., Julien J.F., Julliard R., Machon N. & Le Viol I.
    2013
    Insect Conservation and Diversity
    http://onlineli brary.wiley.com /doi/10.1111/j. 1752-4598.2012. 00217.x/abst...
    Yes

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    Human Activities in Natura 2000 Sites: A Highly Diversified Conservation Network
    Tsiafouli, M.A., Apostolopoulou, E., Mazaris, A.D., Kallimanis A.S., Drakou, E.G. Pantis, J.D.
    2013
    Environmental Management
    http://link.spr inger.com/artic le/10.1007%2Fs0 0267-013-0036-6
    Yes

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    Accounting for the capacity of common and rare species to contribute to diversity spatial patterns: is it a sampling issue or a biological effect?
    Mazaris, A.D., Tsianou, M.A., Sigkounas, A., Dimopoulos, P., Pantis, J.D., Sgardelis, S.P., Kallimanis, A.S.
    2013
    Ecological Indicators
    http://www.scie ncedirect.com/s cience/article/ pii/S1470160X13 000976
    Yes

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    Governance rescaling and the neoliberalization of nature conservation in the EU: We are not all in this together
    Apostolopoulou E., Bormpoudakis D., Paloniemi R., Cent J., Grodzińska-Jurczak M., Pietrzyk-Kaszyńska A., Pantis JD.

    International Journal of Sustainable Development and World Ecology

    Yes

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    Taxon-and-Area Invariances, Maximum Entropy, and the Species-Area Relationship: Reply to Harte et al.
    Šizling, A.L., Kunin, W. & Storch, D.
    2013
    American Naturalist
    http://www.jsto r.org/stable/10 .1086/668822
    Yes

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    Scale sensitivity of drivers of environmental change across Europe
    Tzanopoulos, J., Mouttet, R., Letourneau, A., Vogiatzakis, I.N., Potts, S.G., Henle, K., Mathevet, R., Marty, P.
    2013
    Global Environmental Change
    http://www.scie ncedirect.com/s cience/article/ pii/S0959378012 001124
    Yes

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    More and more generalists: two decades of changes in the European avifauna
    PDF download
    Le Viol I. Jiguet F., Brotons L., Herrando S., Lindström A., Pearce-Higgins J, Reif J., Van Turnhout C. & Devictor V.
    2012
    Biology Letters
    http://rsbl.roy alsocietypublis hing.org/conten t/early/2012/07 /11/rsbl.201...
    Yes
    206.7KB

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    Using species combinations in indicator value analyses
    PDF download
    Cáceres M., Legendre P., Wiser S.K., Brotons L.
    2012
    Methods in Ecology and Evolution
    http://www.scal es-project.net/ page.php?P=37&S P=72
    Yes
    495.92KB

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    Assessing the Natura 2000 network with a common breeding birds survey
    Pellissier V, Touroult J, Julliard R, Siblet J-P, Jiguet F.
    2013
    Animal Conservation
    http://onlineli brary.wiley.com /doi/10.1111/ac v.12030/abstrac t
    Yes

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    Species’ thermal preferences affect forest bird communities along landscape and local scale habitat gradients
    Barnagaud J-Y, Barbaro L, Hampe A, Jiguet F, Archaux F.
    2013
    Ecography
    http://onlineli brary.wiley.com /doi/10.1111/j. 1600-0587.2012. 00227.x/abst...
    Yes

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    Mapping functional connectivity: a landscape approach for handling multiple ecological requirements
    Mimet A, Houet T, Julliard R, Simon L.
    2013
    Methods in Ecology and Evolution
    http://onlineli brary.wiley.com /doi/10.1111/20 41-210x.12024/a bstract
    Yes

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    The whereabouts of flower visitors: contrasting land-use preferences revealed by a country-wide survey based on citizen science
    PDF download
    Deguines N, Julliard R, De Flores M, Fontaine C.
    2012
    PLoS ONE
    http://www.plos one.org/article /info%3Adoi%2F1 0.1371%2Fjourna l.pone.0045822
    Yes
    513.33KB

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    French citizens monitoring ordinary birds provide tools for conservation and ecological sciences
    Jiguet F., Devictor V., Julliard R., Couvet D.
    2012
    Acta Oecologica
    http://www.scie ncedirect.com/s cience/article/ pii/S1146609X11 000762
    Yes

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    Uncertainty in thermal tolerances and climatic debt
    Devictor V., van Swaay C., Brereton T., Brotons L., Chamberlain D., Heliölä J., Herrando S., Julliard R., Kuussaari M., Lindström Å., Reif J., Roy D.B., Schweiger O., Settele J., Stefanescu C., Van Strien A., Van Turnhout C., Vermouzek Z., Wallis De Vries M., Wynhoff I., Jiguet F.
    2012
    Nature Climate Change
    http://www.natu re.com/nclimate /journal/v2/n9/ full/nclimate16 67.html#access
    Yes

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    A review on spatial data integration across monitoring schemes: mapping biodiversity from multiple monitoring data sources
    Sardà-Palomera F., Brotons L.
    2013


    Yes

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    Downscaling of supra-national heterogeneous coarse atlas data to obtain fine-grained distribution maps
    Sardà-Palomera F., Brotons L.
    2013


    Yes

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    Nature Conservation – a new dimension in Open Access publishing bridging science and application
    PDF download
    Henle K., Bell S., Brotons L., Clobert J., Evans D., Goerg Ch., Grodzinska-Jurczak M., Gruber B., Haila Y., Henry P., Huth A., Julliard R., Keil P., Kleyer M., Kotze J., Kunin W., Lengyel Sz., Lin Y., Loyau A., Luck G., Magnuson W., Margules Ch., Matsinos Y., May P., Sousa-Pinto I., Possingham H., Potts S., Ring I., Pryke J., Samways M., Saunders D., Schmeller D., Simila J., Sommer S., Steffan-Dewenter I., Stoev P., Sykes M., Tóthmérész B., Yam R., Tzanopoulos J., Penev L.
    2012
    Nature Conservation
    http://www.pens oft.net/journal s/natureconserv ation/article/3 081/abstract/
    Yes
    983.49KB

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    Factors influencing perception of protected areas – the case of Natura 2000 in Polish Carpathian communities
    Pietrzyk-Kaszyńska A., Cent J., Grodzińska-Jurczak M., Szymańska M.
    2012
    Journal for Nature Conservation
    http://www.scie ncedirect.com/s cience/article/ pii/S1617138112 000556
    Yes

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    Problemy społeczno-ekonomiczne przy wyznaczaniu obszarów Natura 2000 w Polsce. Socio-economic problems during Natura 2000 site selection process
    PDF download
    Grodzińska-Jurczak M., Boćkowski M., Cent J., Pietrzyk-Kaszyńska A.

    Teka Kom. Ochr. Kszt. Środ. Przyr. OL PAN w Lublinie
    http://www.pan- ol.lublin.pl/wy dawnictwa/TOchr 9/Grodzinska.pd f
    Yes
    140.88KB

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    Biodiversity conservation across scales: lessons from a science–policy dialogue
    PDF download
    Paloniemi R, Apostolopoulou E, Primmer E, Grodzinska-Jurcak M, Henle K, Ring I, Kettunen M, Tzanopoulos J, Potts S, van den Hove S, Marty P, McConville A, Simila J
    2012
    Nature Conservation
    http://www.pens oft.net/journal s/natureconserv ation/article/3 144/biodiver...
    Yes
    1.31MB

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    Deliberation in cooperative networks for forest conservation
    Borg, R., Paloniemi, R.
    2012
    Journal of Integrative Environmental Sciences
    http://www.tand fonline.com/doi /abs/10.1080/19 43815X.2012.709 869?journalC...
    Yes

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    Frames of Scale Challenges in Finnish and Greek Biodiversity Conservation
    PDF download
    Apostolopoulou E., Paloniemi R.
    2012
    Ecology and Society
    http://www.ecol ogyandsociety.o rg/vol17/iss4/a rt9/
    Yes
    591.44KB

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    Participation in the management of Greek Natura 2000 sites: Evidence from a cross-level analysis
    Apostolopoulou, E., Drakou, E.G., Pediaditi, K.
    2012
    Journal of Environmental Management
    http://www.ncbi .nlm.nih.gov/pu bmed/23072869
    Yes

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    Mapping from heterogeneous biodiversity monitoring data sources
    Francesc Sardà-Palomera, Lluís Brotons, Dani Villero, Henk Sierdsema, Stuart E. Newson & Frédéric Jiguet
    2012
    Biodiversity and Conservation
    http://link.spr inger.com/artic le/10.1007%2Fs1 0531-012-0347-6 #page-1
    Yes

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    Modelling invasive alien species distributions from digital biodiversity atlases. Model upscaling as a means of reconciling data at different scales
    Arnald Marcer, Joan Pino, Xavier Pons and Lluís Brotons
    2012
    Diversity and Distributions
    http://onlineli brary.wiley.com /doi/10.1111/j. 1472-4642.2012. 00911.x/abst...
    Yes

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    Priorities for biodiversity monitoring in Europe: a review of supranational policies and a novel scheme for integrative prioritization

    Henle, K., B. Bauch, M. Auliya, M. Külvik, G. Pe'er, D. S. Schmeller, Framstad E.
    2013
    Ecological Indicators
    http://www.scie ncedirect.com/s cience/article/ pii/S1470160X13 001416
    Yes

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    A protocol for better design, application and communication of population viability analyses
    Pe’er, G., Matsinos Y. G., Johst K., Franz K. W., Turlure C., Radchuk V., Malinowska A. H., Curtis J. M. R., Naujokaitis-Lewis I., Wintle B. A., Henle K.
    2013
    Conservation Biology
    644-656
    Yes

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    Projecting species’ range expansion dynamics: sources of systematic biases when scaling up patterns and processes
    PDF download
    Bocedi, G., Pe’er G., Heikkinen R.K., Matsinos Y.G., Travis J.M.J.
    2012
    Methods in Ecology and Evolution
    http://onlineli brary.wiley.com /doi/10.1111/j. 2041-210X.2012. 00235.x/abst...
    Yes
    7.38MB

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    Calibration of hybrid species distribution models: the value of general-purpose vs. targeted monitoring data
    De Cáceres, M., Brotons, L
    2012
    Diversity and Distributions
    http://onlineli brary.wiley.com /doi/10.1111/j. 1472-4642.2012. 00899.x/abst...
    Yes

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    Geographical variability in propagule pressure and climatic suitability explain the European distribution of two highly invasive crayfish
    César Capinha, Lluís Brotons and Pedro Anastácio
    2012
    Journal of Biogeography
    http://onlineli brary.wiley.com /doi/10.1111/jb i.12025/abstrac t
    Yes

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    Modeling bird species distribution change in fire prone Mediterranean landscapes: incorporating species dispersal and landscape dynamics
    Lluís Brotons, Miquel De Cáceres, Andrew Fall and Marie-Josée Fortin
    2012
    Ecography
    http://onlineli brary.wiley.com /doi/10.1111/ec og.2012.35.issu e-5/issuetoc
    Yes

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    Influence of Admixture and Paleolithic Range Contractions on Current European Diversity Gradients
    PDF download
    Miguel Arenas
    2013
    Molecular Biology and Evolution
    http://mbe.oxfo rdjournals.org/ content/30/1/57
    Yes
    1.99MB

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    Grassland diversity under changing productivity and the underlying mechanisms – results of a 10-yr experiment
    Liira J., Ingerpuu N., Kalamees R., Moora M., Pärtel M, Püssa K., Roosaluste E., Saar L., Tamme R., Zobel K., Zobel M.
    2012
    Journal of Vegetation Science
    http://onlineli brary.wiley.com /doi/10.1111/j. 1654-1103.2012. 01409.x/abst...
    Yes

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    Does the interpolation accuracy of species distribution models come at the expense of transferability?
    Heikkinen, R.K., Marmion, M. & Luoto, M.
    2012
    Ecography
    http://onlineli brary.wiley.com /doi/10.1111/j. 1600-0587.2011. 06999.x/abst...
    Yes

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    Differences in the climatic debts of birds and butterflies at a continental scale
    evictor, V., van Swaay, C., Brereton, T., Brotons, L., Chamberlain, D., Heliölä, J., Herrando, S., Julliard, R., Kuussaari, M., Lindström, Å., Reif, J., Roy, D., van Strien, A., Settele, J., Schweiger, O., Stefanescu, C., Vermouzek, Z., van Turnhout, C., Wallis de Vries, M. & Jiguet, F.
    2012
    Nature Climate Change
    http://www.natu re.com/nclimate /journal/v2/n2/ full/nclimate13 47.html
    Yes

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    Landscape context affects the relationship between local and landscape species richness of butterflies in semi-natural habitats
    Ekroos, J. & Kuussaari, M.
    2013
    Ecography
    http://onlineli brary.wiley.com /doi/10.1111/j. 1600-0587.2011. 07005.x/abst...
    Yes

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    Effect of habitat area and isolation on plant trait distribution in European forests and grasslands
    Lindborg, R., Helm, A., Bommarco, R., Heikkinen, R.K., Kühn, I., Pykälä, J. & Pärtel, M.
    2012
    Ecography
    http://onlineli brary.wiley.com /doi/10.1111/j. 1600-0587.2011. 07286.x/abst...
    Yes

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    The landscape matrix modifies the effect of habitat fragmentation in grassland butterflies
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